SOLTUB.AGRIA.G00000009645

Description : Alcohol dehydrogenase

Gene families : OG_02_0001222 (Orthogroups with 8 Potato genotypes) Phylogenetic Tree(s): No tree available for this family

Sequence : coding (download), protein (download)

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Note:Only the main profile, including all conditions, is shown. Additional statistics and tissue specific profiles are available here.

Type	Description	Actions
Neighborhood	HRR Network (HRR 50 & PCC 0.5): SOLTUB.AGRIA.G00000009645
Cluster	HCCA Clusters (HRR 50 & PCC 0.5): Cluster_121

Type	GO Term	Name	Evidence	Source
BP	GO:0001523	retinoid metabolic process	IEA	AHRD
BP	GO:0002237	response to molecule of bacterial origin	None	Extended
BP	GO:0003008	system process	None	Extended
BP	GO:0003016	respiratory system process	IEA	AHRD
MF	GO:0003674	molecular_function	None	Extended
MF	GO:0003824	catalytic activity	None	Extended
MF	GO:0004022	alcohol dehydrogenase (NAD+) activity	IEA	AHRD
MF	GO:0004552	octanol dehydrogenase activity	IEA	AHRD
MF	GO:0005488	binding	None	Extended
MF	GO:0005515	protein binding	None	Extended
CC	GO:0005575	cellular_component	None	Extended
CC	GO:0005634	nucleus	IEA	AHRD
CC	GO:0005739	mitochondrion	IEA	AHRD
CC	GO:0005829	cytosol	IEA	AHRD
BP	GO:0006066	alcohol metabolic process	IEA	AHRD
BP	GO:0006081	cellular aldehyde metabolic process	None	Extended
BP	GO:0006629	lipid metabolic process	None	Extended
BP	GO:0006720	isoprenoid metabolic process	None	Extended
BP	GO:0006721	terpenoid metabolic process	None	Extended
BP	GO:0006807	nitrogen compound metabolic process	None	Extended
BP	GO:0006950	response to stress	None	Extended
BP	GO:0007610	behavior	None	Extended
BP	GO:0007611	learning or memory	None	Extended
BP	GO:0007612	learning	None	Extended
BP	GO:0007632	visual behavior	None	Extended
BP	GO:0008150	biological_process	None	Extended
BP	GO:0008152	metabolic process	None	Extended
BP	GO:0008217	regulation of blood pressure	None	Extended
BP	GO:0008299	isoprenoid biosynthetic process	IEA	AHRD
BP	GO:0008306	associative learning	None	Extended
BP	GO:0008542	visual learning	IEA	AHRD
BP	GO:0008610	lipid biosynthetic process	None	Extended
BP	GO:0009056	catabolic process	None	Extended
BP	GO:0009058	biosynthetic process	None	Extended
BP	GO:0009314	response to radiation	None	Extended
BP	GO:0009416	response to light stimulus	None	Extended
BP	GO:0009605	response to external stimulus	None	Extended
BP	GO:0009607	response to biotic stimulus	None	Extended
BP	GO:0009628	response to abiotic stimulus	None	Extended
BP	GO:0009719	response to endogenous stimulus	None	Extended
BP	GO:0009725	response to hormone	None	Extended
BP	GO:0009753	response to jasmonic acid	IEA	AHRD
BP	GO:0009820	alkaloid metabolic process	None	Extended
BP	GO:0009987	cellular process	None	Extended
BP	GO:0010033	response to organic substance	None	Extended
BP	GO:0016101	diterpenoid metabolic process	None	Extended
MF	GO:0016491	oxidoreductase activity	None	Extended
MF	GO:0016614	oxidoreductase activity, acting on CH-OH group of donors	None	Extended
MF	GO:0016616	oxidoreductase activity, acting on the CH-OH group of donors, NAD or NADP as acceptor	None	Extended
MF	GO:0016651	oxidoreductase activity, acting on NAD(P)H	None	Extended
MF	GO:0016657	oxidoreductase activity, acting on NAD(P)H, nitrogenous group as acceptor	None	Extended
MF	GO:0018455	alcohol dehydrogenase [NAD(P)+] activity	None	Extended
BP	GO:0019222	regulation of metabolic process	None	Extended
BP	GO:0031399	regulation of protein modification process	None	Extended
BP	GO:0032496	response to lipopolysaccharide	IEA	AHRD
BP	GO:0032501	multicellular organismal process	None	Extended
BP	GO:0033993	response to lipid	None	Extended
BP	GO:0035834	indole alkaloid metabolic process	IEA	AHRD
BP	GO:0042221	response to chemical	None	Extended
MF	GO:0042802	identical protein binding	IEA	AHRD
MF	GO:0043167	ion binding	None	Extended
MF	GO:0043169	cation binding	None	Extended
BP	GO:0043207	response to external biotic stimulus	None	Extended
CC	GO:0043226	organelle	None	Extended
CC	GO:0043227	membrane-bounded organelle	None	Extended
CC	GO:0043229	intracellular organelle	None	Extended
CC	GO:0043231	intracellular membrane-bounded organelle	None	Extended
BP	GO:0044237	cellular metabolic process	None	Extended
BP	GO:0044238	primary metabolic process	None	Extended
BP	GO:0044248	cellular catabolic process	None	Extended
BP	GO:0044249	cellular biosynthetic process	None	Extended
BP	GO:0044255	cellular lipid metabolic process	None	Extended
BP	GO:0044281	small molecule metabolic process	None	Extended
BP	GO:0044282	small molecule catabolic process	None	Extended
BP	GO:0045777	positive regulation of blood pressure	IEA	AHRD
BP	GO:0046185	aldehyde catabolic process	None	Extended
BP	GO:0046292	formaldehyde metabolic process	None	Extended
BP	GO:0046294	formaldehyde catabolic process	IEA	AHRD
MF	GO:0046872	metal ion binding	IEA	AHRD
BP	GO:0050789	regulation of biological process	None	Extended
BP	GO:0050877	nervous system process	None	Extended
BP	GO:0050890	cognition	None	Extended
BP	GO:0050896	response to stimulus	None	Extended
BP	GO:0051171	regulation of nitrogen compound metabolic process	None	Extended
BP	GO:0051246	regulation of protein metabolic process	None	Extended
BP	GO:0051409	response to nitrosative stress	IEA	AHRD
MF	GO:0051903	S-(hydroxymethyl)glutathione dehydrogenase activity	IEA	AHRD
BP	GO:0060255	regulation of macromolecule metabolic process	None	Extended
BP	GO:0065007	biological regulation	None	Extended
BP	GO:0065008	regulation of biological quality	None	Extended
BP	GO:0070542	response to fatty acid	None	Extended
BP	GO:0071704	organic substance metabolic process	None	Extended
MF	GO:0080007	S-nitrosoglutathione reductase activity	IEA	AHRD
BP	GO:0080090	regulation of primary metabolic process	None	Extended
BP	GO:0098754	detoxification	None	Extended
MF	GO:0102311	8-hydroxygeraniol dehydrogenase activity	IEA	AHRD
BP	GO:0110095	cellular detoxification of aldehyde	None	Extended
CC	GO:0110165	cellular anatomical entity	None	Extended
BP	GO:1901564	organonitrogen compound metabolic process	None	Extended
BP	GO:1901575	organic substance catabolic process	None	Extended
BP	GO:1901576	organic substance biosynthetic process	None	Extended
BP	GO:1901615	organic hydroxy compound metabolic process	None	Extended
BP	GO:1901700	response to oxygen-containing compound	None	Extended
BP	GO:1990748	cellular detoxification	None	Extended
BP	GO:2000169	regulation of peptidyl-cysteine S-nitrosylation	IEA	AHRD

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Type	GO Term	Name	Evidence	Source
MF	GO:0000293	ferric-chelate reductase activity	IEP	Predict GO terms from Neighborhoods
BP	GO:0000578	embryonic axis specification	IEP	Predict GO terms from Neighborhoods
BP	GO:0000902	cell morphogenesis	IEP	Predict GO terms from Neighborhoods
BP	GO:0000904	cell morphogenesis involved in differentiation	IEP	Predict GO terms from Neighborhoods
BP	GO:0003006	developmental process involved in reproduction	IEP	Predict GO terms from Neighborhoods
MF	GO:0004175	endopeptidase activity	IEP	Predict GO terms from Neighborhoods
MF	GO:0004252	serine-type endopeptidase activity	IEP	Predict GO terms from Neighborhoods
MF	GO:0005262	calcium channel activity	IEP	Predict GO terms from Neighborhoods
CC	GO:0005615	extracellular space	IEP	Predict GO terms from Neighborhoods
CC	GO:0005618	cell wall	IEP	Predict GO terms from Neighborhoods
CC	GO:0005783	endoplasmic reticulum	IEP	Predict GO terms from Neighborhoods
CC	GO:0005874	microtubule	IEP	Predict GO terms from Neighborhoods
CC	GO:0005881	cytoplasmic microtubule	IEP	Predict GO terms from Neighborhoods
BP	GO:0006012	galactose metabolic process	IEP	Predict GO terms from Neighborhoods
BP	GO:0006508	proteolysis	IEP	Predict GO terms from Neighborhoods
BP	GO:0006714	sesquiterpenoid metabolic process	IEP	Predict GO terms from Neighborhoods
BP	GO:0006801	superoxide metabolic process	IEP	Predict GO terms from Neighborhoods
BP	GO:0006879	cellular iron ion homeostasis	IEP	Predict GO terms from Neighborhoods
MF	GO:0008157	protein phosphatase 1 binding	IEP	Predict GO terms from Neighborhoods
MF	GO:0008233	peptidase activity	IEP	Predict GO terms from Neighborhoods
MF	GO:0008236	serine-type peptidase activity	IEP	Predict GO terms from Neighborhoods
MF	GO:0008378	galactosyltransferase activity	IEP	Predict GO terms from Neighborhoods
MF	GO:0008381	mechanosensitive ion channel activity	IEP	Predict GO terms from Neighborhoods
BP	GO:0009312	oligosaccharide biosynthetic process	IEP	Predict GO terms from Neighborhoods
CC	GO:0009505	plant-type cell wall	IEP	Predict GO terms from Neighborhoods
CC	GO:0009506	plasmodesma	IEP	Predict GO terms from Neighborhoods
BP	GO:0009566	fertilization	IEP	Predict GO terms from Neighborhoods
BP	GO:0009567	double fertilization forming a zygote and endosperm	IEP	Predict GO terms from Neighborhoods
BP	GO:0009606	tropism	IEP	Predict GO terms from Neighborhoods
BP	GO:0009617	response to bacterium	IEP	Predict GO terms from Neighborhoods
BP	GO:0009629	response to gravity	IEP	Predict GO terms from Neighborhoods
BP	GO:0009630	gravitropism	IEP	Predict GO terms from Neighborhoods
BP	GO:0009653	anatomical structure morphogenesis	IEP	Predict GO terms from Neighborhoods
BP	GO:0009664	plant-type cell wall organization	IEP	Predict GO terms from Neighborhoods
BP	GO:0009682	induced systemic resistance	IEP	Predict GO terms from Neighborhoods
BP	GO:0009704	de-etiolation	IEP	Predict GO terms from Neighborhoods
BP	GO:0009733	response to auxin	IEP	Predict GO terms from Neighborhoods
BP	GO:0009826	unidimensional cell growth	IEP	Predict GO terms from Neighborhoods
BP	GO:0009827	plant-type cell wall modification	IEP	Predict GO terms from Neighborhoods
BP	GO:0009828	plant-type cell wall loosening	IEP	Predict GO terms from Neighborhoods
BP	GO:0009860	pollen tube growth	IEP	Predict GO terms from Neighborhoods
BP	GO:0009913	epidermal cell differentiation	IEP	Predict GO terms from Neighborhoods
CC	GO:0009921	auxin efflux carrier complex	IEP	Predict GO terms from Neighborhoods
CC	GO:0009925	basal plasma membrane	IEP	Predict GO terms from Neighborhoods
BP	GO:0009932	cell tip growth	IEP	Predict GO terms from Neighborhoods
BP	GO:0009942	longitudinal axis specification	IEP	Predict GO terms from Neighborhoods
BP	GO:0009959	negative gravitropism	IEP	Predict GO terms from Neighborhoods
CC	GO:0010005	cortical microtubule, transverse to long axis	IEP	Predict GO terms from Neighborhoods
BP	GO:0010015	root morphogenesis	IEP	Predict GO terms from Neighborhoods
BP	GO:0010029	regulation of seed germination	IEP	Predict GO terms from Neighborhoods
BP	GO:0010051	xylem and phloem pattern formation	IEP	Predict GO terms from Neighborhoods
BP	GO:0010074	maintenance of meristem identity	IEP	Predict GO terms from Neighborhoods
BP	GO:0010101	post-embryonic root morphogenesis	IEP	Predict GO terms from Neighborhoods
BP	GO:0010102	lateral root morphogenesis	IEP	Predict GO terms from Neighborhoods
BP	GO:0010103	stomatal complex morphogenesis	IEP	Predict GO terms from Neighborhoods
BP	GO:0010150	leaf senescence	IEP	Predict GO terms from Neighborhoods
BP	GO:0010315	auxin export across the plasma membrane	IEP	Predict GO terms from Neighborhoods
BP	GO:0010325	raffinose family oligosaccharide biosynthetic process	IEP	Predict GO terms from Neighborhoods
MF	GO:0010329	auxin efflux transmembrane transporter activity	IEP	Predict GO terms from Neighborhoods
MF	GO:0010334	sesquiterpene synthase activity	IEP	Predict GO terms from Neighborhoods
BP	GO:0010338	leaf formation	IEP	Predict GO terms from Neighborhoods
BP	GO:0010358	leaf shaping	IEP	Predict GO terms from Neighborhoods
BP	GO:0015772	oligosaccharide transport	IEP	Predict GO terms from Neighborhoods
BP	GO:0015773	raffinose transport	IEP	Predict GO terms from Neighborhoods
BP	GO:0016049	cell growth	IEP	Predict GO terms from Neighborhoods
BP	GO:0016106	sesquiterpenoid biosynthetic process	IEP	Predict GO terms from Neighborhoods
MF	GO:0016174	NAD(P)H oxidase H2O2-forming activity	IEP	Predict GO terms from Neighborhoods
MF	GO:0016722	oxidoreductase activity, acting on metal ions	IEP	Predict GO terms from Neighborhoods
MF	GO:0017171	serine hydrolase activity	IEP	Predict GO terms from Neighborhoods
MF	GO:0019208	phosphatase regulator activity	IEP	Predict GO terms from Neighborhoods
BP	GO:0019827	stem cell population maintenance	IEP	Predict GO terms from Neighborhoods
MF	GO:0019888	protein phosphatase regulator activity	IEP	Predict GO terms from Neighborhoods
BP	GO:0022414	reproductive process	IEP	Predict GO terms from Neighborhoods
BP	GO:0030001	metal ion transport	IEP	Predict GO terms from Neighborhoods
BP	GO:0030177	positive regulation of Wnt signaling pathway	IEP	Predict GO terms from Neighborhoods
BP	GO:0030178	negative regulation of Wnt signaling pathway	IEP	Predict GO terms from Neighborhoods
CC	GO:0030312	external encapsulating structure	IEP	Predict GO terms from Neighborhoods
CC	GO:0031012	extracellular matrix	IEP	Predict GO terms from Neighborhoods
BP	GO:0032502	developmental process	IEP	Predict GO terms from Neighborhoods
BP	GO:0032874	positive regulation of stress-activated MAPK cascade	IEP	Predict GO terms from Neighborhoods
BP	GO:0033212	iron import into cell	IEP	Predict GO terms from Neighborhoods
BP	GO:0033214	siderophore-dependent iron import into cell	IEP	Predict GO terms from Neighborhoods
BP	GO:0033383	geranyl diphosphate metabolic process	IEP	Predict GO terms from Neighborhoods
MF	GO:0035250	UDP-galactosyltransferase activity	IEP	Predict GO terms from Neighborhoods
BP	GO:0035304	regulation of protein dephosphorylation	IEP	Predict GO terms from Neighborhoods
BP	GO:0036473	cell death in response to oxidative stress	IEP	Predict GO terms from Neighborhoods
BP	GO:0040007	growth	IEP	Predict GO terms from Neighborhoods
BP	GO:0042545	cell wall modification	IEP	Predict GO terms from Neighborhoods
BP	GO:0043171	peptide catabolic process	IEP	Predict GO terms from Neighborhoods
BP	GO:0043476	pigment accumulation	IEP	Predict GO terms from Neighborhoods
BP	GO:0043478	pigment accumulation in response to UV light	IEP	Predict GO terms from Neighborhoods
BP	GO:0043479	pigment accumulation in tissues in response to UV light	IEP	Predict GO terms from Neighborhoods
BP	GO:0043480	pigment accumulation in tissues	IEP	Predict GO terms from Neighborhoods
BP	GO:0043481	anthocyanin accumulation in tissues in response to UV light	IEP	Predict GO terms from Neighborhoods
CC	GO:0045177	apical part of cell	IEP	Predict GO terms from Neighborhoods
BP	GO:0045338	farnesyl diphosphate metabolic process	IEP	Predict GO terms from Neighborhoods
BP	GO:0046328	regulation of JNK cascade	IEP	Predict GO terms from Neighborhoods
BP	GO:0046330	positive regulation of JNK cascade	IEP	Predict GO terms from Neighborhoods
MF	GO:0047216	inositol 3-alpha-galactosyltransferase activity	IEP	Predict GO terms from Neighborhoods
MF	GO:0047461	(+)-delta-cadinene synthase activity	IEP	Predict GO terms from Neighborhoods
BP	GO:0048366	leaf development	IEP	Predict GO terms from Neighborhoods
BP	GO:0048442	sepal development	IEP	Predict GO terms from Neighborhoods
BP	GO:0048480	stigma development	IEP	Predict GO terms from Neighborhoods
BP	GO:0048497	maintenance of floral organ identity	IEP	Predict GO terms from Neighborhoods
BP	GO:0048580	regulation of post-embryonic development	IEP	Predict GO terms from Neighborhoods
BP	GO:0048581	negative regulation of post-embryonic development	IEP	Predict GO terms from Neighborhoods
BP	GO:0048588	developmental cell growth	IEP	Predict GO terms from Neighborhoods
BP	GO:0048589	developmental growth	IEP	Predict GO terms from Neighborhoods
BP	GO:0048764	trichoblast maturation	IEP	Predict GO terms from Neighborhoods
BP	GO:0048765	root hair cell differentiation	IEP	Predict GO terms from Neighborhoods
BP	GO:0048825	cotyledon development	IEP	Predict GO terms from Neighborhoods
BP	GO:0048826	cotyledon morphogenesis	IEP	Predict GO terms from Neighborhoods
BP	GO:0048827	phyllome development	IEP	Predict GO terms from Neighborhoods
BP	GO:0048830	adventitious root development	IEP	Predict GO terms from Neighborhoods
MF	GO:0050664	oxidoreductase activity, acting on NAD(P)H, oxygen as acceptor	IEP	Predict GO terms from Neighborhoods
BP	GO:0050793	regulation of developmental process	IEP	Predict GO terms from Neighborhoods
BP	GO:0050982	detection of mechanical stimulus	IEP	Predict GO terms from Neighborhoods
BP	GO:0051093	negative regulation of developmental process	IEP	Predict GO terms from Neighborhoods
BP	GO:0051239	regulation of multicellular organismal process	IEP	Predict GO terms from Neighborhoods
BP	GO:0051241	negative regulation of multicellular organismal process	IEP	Predict GO terms from Neighborhoods
BP	GO:0052575	carbohydrate localization	IEP	Predict GO terms from Neighborhoods
BP	GO:0052576	carbohydrate storage	IEP	Predict GO terms from Neighborhoods
CC	GO:0055028	cortical microtubule	IEP	Predict GO terms from Neighborhoods
BP	GO:0060560	developmental growth involved in morphogenesis	IEP	Predict GO terms from Neighborhoods
BP	GO:0070304	positive regulation of stress-activated protein kinase signaling cascade	IEP	Predict GO terms from Neighborhoods
BP	GO:0070482	response to oxygen levels	IEP	Predict GO terms from Neighborhoods
BP	GO:0071465	cellular response to desiccation	IEP	Predict GO terms from Neighborhoods
BP	GO:0071554	cell wall organization or biogenesis	IEP	Predict GO terms from Neighborhoods
BP	GO:0071669	plant-type cell wall organization or biogenesis	IEP	Predict GO terms from Neighborhoods
MF	GO:0080161	auxin transmembrane transporter activity	IEP	Predict GO terms from Neighborhoods
BP	GO:0080162	endoplasmic reticulum to cytosol auxin transport	IEP	Predict GO terms from Neighborhoods
BP	GO:0090090	negative regulation of canonical Wnt signaling pathway	IEP	Predict GO terms from Neighborhoods
BP	GO:0090175	regulation of establishment of planar polarity	IEP	Predict GO terms from Neighborhoods
BP	GO:0090377	seed trichome initiation	IEP	Predict GO terms from Neighborhoods
BP	GO:0090626	plant epidermis morphogenesis	IEP	Predict GO terms from Neighborhoods
BP	GO:0090693	plant organ senescence	IEP	Predict GO terms from Neighborhoods
BP	GO:0090697	post-embryonic plant organ morphogenesis	IEP	Predict GO terms from Neighborhoods
BP	GO:0090698	post-embryonic plant morphogenesis	IEP	Predict GO terms from Neighborhoods
BP	GO:0090700	maintenance of plant organ identity	IEP	Predict GO terms from Neighborhoods
BP	GO:0097264	self proteolysis	IEP	Predict GO terms from Neighborhoods
BP	GO:0097468	programmed cell death in response to reactive oxygen species	IEP	Predict GO terms from Neighborhoods
BP	GO:0098727	maintenance of cell number	IEP	Predict GO terms from Neighborhoods
CC	GO:0099081	supramolecular polymer	IEP	Predict GO terms from Neighborhoods
CC	GO:0099512	supramolecular fiber	IEP	Predict GO terms from Neighborhoods
CC	GO:0099513	polymeric cytoskeletal fiber	IEP	Predict GO terms from Neighborhoods
BP	GO:0120251	hydrocarbon biosynthetic process	IEP	Predict GO terms from Neighborhoods
BP	GO:0120252	hydrocarbon metabolic process	IEP	Predict GO terms from Neighborhoods
BP	GO:1900140	regulation of seedling development	IEP	Predict GO terms from Neighborhoods
BP	GO:1900367	positive regulation of defense response to insect	IEP	Predict GO terms from Neighborhoods
BP	GO:1901332	negative regulation of lateral root development	IEP	Predict GO terms from Neighborhoods
BP	GO:1901678	iron coordination entity transport	IEP	Predict GO terms from Neighborhoods
BP	GO:1901926	cadinene metabolic process	IEP	Predict GO terms from Neighborhoods
BP	GO:1901928	cadinene biosynthetic process	IEP	Predict GO terms from Neighborhoods
BP	GO:1902066	regulation of cell wall pectin metabolic process	IEP	Predict GO terms from Neighborhoods
BP	GO:1902531	regulation of intracellular signal transduction	IEP	Predict GO terms from Neighborhoods
BP	GO:1902533	positive regulation of intracellular signal transduction	IEP	Predict GO terms from Neighborhoods
BP	GO:1905392	plant organ morphogenesis	IEP	Predict GO terms from Neighborhoods
BP	GO:2000026	regulation of multicellular organismal development	IEP	Predict GO terms from Neighborhoods
BP	GO:2000027	regulation of animal organ morphogenesis	IEP	Predict GO terms from Neighborhoods
BP	GO:2000033	regulation of seed dormancy process	IEP	Predict GO terms from Neighborhoods
BP	GO:2000034	regulation of seed maturation	IEP	Predict GO terms from Neighborhoods
BP	GO:2000050	regulation of non-canonical Wnt signaling pathway	IEP	Predict GO terms from Neighborhoods
BP	GO:2000052	positive regulation of non-canonical Wnt signaling pathway	IEP	Predict GO terms from Neighborhoods
BP	GO:2000095	regulation of Wnt signaling pathway, planar cell polarity pathway	IEP	Predict GO terms from Neighborhoods
BP	GO:2000096	positive regulation of Wnt signaling pathway, planar cell polarity pathway	IEP	Predict GO terms from Neighborhoods
BP	GO:2000122	negative regulation of stomatal complex development	IEP	Predict GO terms from Neighborhoods
BP	GO:2001088	trisaccharide transport	IEP	Predict GO terms from Neighborhoods

InterPro domains	Description	Start	Stop
IPR011032	GroES-like_sf	6	181
IPR013149	ADH-like_C	187	318
IPR036291	NAD(P)-bd_dom_sf	148	321
IPR011032	GroES-like_sf	309	358
IPR013154	ADH_N	58	142

External Links

No external refs found!

SOLTUB.AGRIA.G00000009645

Expression Profile

Co-expression Networks

Functional Annotation